Russia has used missiles linked to its S-400 air defence system to attack ground targets in Ukraine, highlighting how Moscow has adapted its air defence weapons for a different role. How is Russia using S-400 missiles for ground attacks? AdvertisementHowever, Russia has previously used similar air defence missiles against targets on the ground. Using air defence missiles for ground attacks can give Russia another way to strike targets when conventional ballistic missiles are in limited supply. India operates the S-400 air defence system, making Russia’s adaptation of the system particularly important for New Delhi.